Note: I also keep (fairly current) publication lists at ORCIDGoogle Scholar, Smithsonian Libraries, and ResearchGate.

In press

S.R. Schachat and P.Z. Goldstein. Macroheterocera (Lepidoptera: Noctuidae: Acronictinae) demonstrate the variable role of wing venation in development of the nymphalid groundplan. Insect Systematics and Diversity: ISD-2017-0054.

Early view

S.R. Schachat and R.L. Brown. Wing patterns of ditrysian moths (Lepidoptera: Psychidae) include variants and violations of predictive models. Austral Entomology: Early view. DOI: 10.1111/aen.12284 [pdf]


S.R. Schachat, C.C. Labandeira, M.R. Saltzman, B.D. Cramer, J.L. Payne, and C.K. Boyce. Phanerozoic pO2 and the early evolution of terrestrial animals. Proceedings of the Royal Society B: Biological Sciences 285(1871): 20172631. DOI: 10.1098/rspb.2017.2631

S.R. Schachat, R.G. Robbins, and J. Goddard. Color Patterning in Hard Ticks (Acari: Ixodidae). Journal of Medical Entomology 55(1): 1-13. DOI: 10.1093/jme/tjx173


S.R. Schachat. Connecting the dots: Spots and bands on the wings of Lichenaula Meyrick, 1890 (Lepidoptera: Xyloryctidae) share a uniform relationship with wing venation. Arthropod Systematics & Phylogeny 75(3): 363-371. [pdf]

S.R. Schachat. The wing pattern of Moerarchis Durrant, 1914 (Lepidoptera: Tineidae) clarifies transitions between predictive models. Royal Society Open Science 4(3). DOI: 10.1098/rsos.161002 [pdf]


S.R. Schachat and G.W. Gibbs. Variable wing venation in Agathiphaga (Lepidoptera: Agathiphagidae) is key to understanding the evolution of basal moths. Royal Society Open Science 3(10). DOI: 10.1098/rsos.160453 [pdf]

S.R. Schachat and R.L. Brown. Forewing color pattern in Micropterigidae (Insecta: Lepidoptera): Homologies between contrast boundaries, and a revised hypothesis for the origin of symmetry systems. BMC Evolutionary Biology 16(116). DOI: 10.1186/s12862-016-0687-z [pdf]

S.R. Schachat, D.G. Mulcahy, and J.R. Mendelson III. Conservation threats and the phylogenetic utility of IUCN Red List rankings in Incilius toads. Conservation Biology 30(1): 72-81. DOI: 10.1111/cobi.12567 [pdf]

S. Ho, S.R. Schachat, W.H. Piel, and A. Monteiro. 2016. Attack risk for butterflies changes with eyespot number and size. Royal Society Open Science 3(1): 150614. DOI: 10.1098/rsos.150614 [pdf]


S.R. Schachat. Insect biodiversity in Meiji and Art Nouveau design. American Entomologist 61(4):215-222. DOI: 10.1093/ae/tmv071 [pdf]

S.R. Schachat, C.C. Labandeira, and D.S. Chaney. Insect herbivory from Early Permian Mitchell Creek Flats of north-central Texas: opportunism in a balanced component community. Palaeogeography, Palaeoclimatology, Palaeoecology 440: 830-847. DOI: 10.1016/j.palaeo.2015.10.001 [pdf]

S.R. Schachat and R.L. Brown. Color pattern on the forewing of Micropterix (Lepidoptera: Micropterigidae): Insights into the evolution of wing pattern and wing venation in moths. PLoS ONE 10(10): e0139972. DOI: 10.1371/journal.pone.0139972 [pdf]

S.R. Schachat and C.C. Labandeira. Evolution of a complex behavior: The origin and initial diversification of foliar galling by Permian insects. The Science of Nature – Naturwissenschaften 102(14): 1-8. DOI: 10.1007/s00114-015-1266-7 [pdf]

S.R. Schachat, J.C. Oliver, and A. Monteiro. Nymphalid eyespots are co-opted to novel wing locations following a similar pattern in independent lineages. BMC Evolutionary Biology 15(20). DOI: 10.1186/s12862-015-0300-x [pdf]


S.R. Schachat. Drawn before Wonderland: Bizarre illustrated insects of the nineteenth century. American Entomologist 60(3): 162-165. DOI: 10.1093/ae/60.3.162 [pdf]

S.R. Schachat, C.C. Labandeira, J. Gordon, D.S. Chaney, S. Levi, M.S. Halthore, and J. Alvarez. Plant-insect interactions from Early Permian (Kungurian) Colwell Creek Pond, north-central Texas: The early spread of herbivory in riparian environments. International Journal of Plant Sciences 175(8): 855-890. DOI: 10.1086/677679 [pdf]

In revision

S.R. Schachat, C.C. Labandeira, and S.A. Maccracken. The importance of sampling standardization for comparisons of insect herbivory in deep time: a case study from the late Paleozoic. Royal Society Open Science: RSOS-171991.