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Note: I also keep (fairly current) publication lists at ORCIDGoogle Scholar, and ResearchGate.


S.R. Schachat and C.C. Labandeira. Are insects heading toward their first mass extinction? Distinguishing turnover from crises in their fossil record. Annals of the Entomological Society of America: Early view. DOI: 10.1093/aesa/saaa042 [pdf]

S.R. Schachat, S.A. Maccracken, and C.C. Labandeira. Sampling fossil floras for the study of insect herbivory: how many leaves is enough? Fossil Record 23(1): 15–32. DOI: 10.5194/fr-23-15-2020 [pdf]

S.R. Schachat. Symmetry systems on the wings of Dichromodes Guenée (Lepidoptera: Geometridae) are unconstrained by venation. PeerJ 8:e8263. DOI: 10.7717/peerj.8263 [pdf]


S.R. Schachat, C.C. Labandeira, M.E. Clapham, and J.L. Payne. A Cretaceous peak in family-level insect diversity estimated with mark–recapture methodology. Proceedings of the Royal Society B: Biological Sciences 286(1917): 20192054. DOI: 10.1098/rspb.2019.2054 [pdf]

S.R. Schachat. The wing pattern of Hydriomena Hübner, [1825] (Lepidoptera: Geometridae: Larentiinae) lacks a predictable relationship with venation. Journal of Morphology 280(11): 1651–1667. DOI: 10.1002/jmor.21055 [pdf]


S.R. Schachat and R.L. Brown. Wing patterns of ditrysian moths (Lepidoptera: Psychidae) include variants and violations of predictive models. Austral Entomology 57(3): 333–343. DOI: 10.1111/aen.12284 [pdf]

S.R. Schachat, C.C. Labandeira, and S.A. Maccracken. The importance of sampling standardization for comparisons of insect herbivory in deep time: a case study from the late Paleozoic. Royal Society Open Science 5(3): 171991. DOI: 10.1098/rsos.171991 [pdf]

S.R. Schachat and P.Z. Goldstein. Acronictinae (Lepidoptera: Noctuidae: Macroheterocera) demonstrate the variable role of wing venation in development of the nymphalid groundplan. Insect Systematics and Diversity 2(2): 4. DOI: 10.1093/isd/ixy004 [pdf]

S.R. Schachat, C.C. Labandeira, M.R. Saltzman, B.D. Cramer, J.L. Payne, and C.K. Boyce. Phanerozoic pO2 and the early evolution of terrestrial animals. Proceedings of the Royal Society B: Biological Sciences 285(1871): 20172631. DOI: 10.1098/rspb.2017.2631 [pdf]

S.R. Schachat, R.G. Robbins, and J. Goddard. Color Patterning in Hard Ticks (Acari: Ixodidae). Journal of Medical Entomology 55(1): 1–13. DOI: 10.1093/jme/tjx173 [pdf]


S.R. Schachat. Connecting the dots: Spots and bands on the wings of Lichenaula Meyrick, 1890 (Lepidoptera: Xyloryctidae) share a uniform relationship with wing venation. Arthropod Systematics & Phylogeny 75(3): 363–371. [pdf]

S.R. Schachat. The wing pattern of Moerarchis Durrant, 1914 (Lepidoptera: Tineidae) clarifies transitions between predictive models. Royal Society Open Science 4(3). DOI: 10.1098/rsos.161002 [pdf]


S.R. Schachat and G.W. Gibbs. Variable wing venation in Agathiphaga (Lepidoptera: Agathiphagidae) is key to understanding the evolution of basal moths. Royal Society Open Science 3(10). DOI: 10.1098/rsos.160453 [pdf]

S.R. Schachat and R.L. Brown. Forewing color pattern in Micropterigidae (Insecta: Lepidoptera): Homologies between contrast boundaries, and a revised hypothesis for the origin of symmetry systems. BMC Evolutionary Biology 16(116). DOI: 10.1186/s12862-016-0687-z [pdf]

S.R. Schachat, D.G. Mulcahy, and J.R. Mendelson III. Conservation threats and the phylogenetic utility of IUCN Red List rankings in Incilius toads. Conservation Biology 30(1): 72–81. DOI: 10.1111/cobi.12567 [pdf]

S. Ho, S.R. Schachat, W.H. Piel, and A. Monteiro. 2016. Attack risk for butterflies changes with eyespot number and size. Royal Society Open Science 3(1): 150614. DOI: 10.1098/rsos.150614 [pdf]


S.R. Schachat. Insect biodiversity in Meiji and Art Nouveau design. American Entomologist 61(4):215–222. DOI: 10.1093/ae/tmv071 [pdf]

S.R. Schachat, C.C. Labandeira, and D.S. Chaney. Insect herbivory from Early Permian Mitchell Creek Flats of north-central Texas: opportunism in a balanced component community. Palaeogeography, Palaeoclimatology, Palaeoecology 440: 830–847. DOI: 10.1016/j.palaeo.2015.10.001 [pdf]

S.R. Schachat and R.L. Brown. Color pattern on the forewing of Micropterix (Lepidoptera: Micropterigidae): Insights into the evolution of wing pattern and wing venation in moths. PLoS ONE 10(10): e0139972. DOI: 10.1371/journal.pone.0139972 [pdf]

S.R. Schachat and C.C. Labandeira. Evolution of a complex behavior: The origin and initial diversification of foliar galling by Permian insects. The Science of Nature – Naturwissenschaften 102(14): 1–8. DOI: 10.1007/s00114-015-1266-7 [pdf]

S.R. Schachat, J.C. Oliver, and A. Monteiro. Nymphalid eyespots are co-opted to novel wing locations following a similar pattern in independent lineages. BMC Evolutionary Biology 15(20). DOI: 10.1186/s12862-015-0300-x [pdf]


S.R. Schachat. Drawn before Wonderland: Bizarre illustrated insects of the nineteenth century. American Entomologist 60(3): 162–165. DOI: 10.1093/ae/60.3.162 [pdf]

S.R. Schachat, C.C. Labandeira, J. Gordon, D.S. Chaney, S. Levi, M.S. Halthore, and J. Alvarez. Plant-insect interactions from Early Permian (Kungurian) Colwell Creek Pond, north-central Texas: The early spread of herbivory in riparian environments. International Journal of Plant Sciences 175(8): 855–890. DOI: 10.1086/677679 [pdf]